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Subcellular Location Features descriptions


2D SLF


Zernike features - A set of circle polynomials in two polar variables: The moments themselves are complex numbers and are sensitive to rotation of the image therefore the magnitudes of the moments were used as features. Zernike moments through degree 12 were used providing a set of 49 rotation invariant features. (Plots of all of the Zernike polynomials can be viewed here).


Haralick texture features - Texture features based in a gray-level co-occurrence matrix of the image: Haralick has described 13 statistics that can be calculated from the co-occurrence matrix of which 13 are used here.

H1. Angular Second Moment
H2. Contrast
H3. Correlation
H4. Sum of Squares:Variation
H5. Inverse Difference Moment
H6. Sum Average
H7. Sum Variance
H8. Sum Entropy
H9. Entropy
H10. Difference VarianceM
H11. Difference Entropy
H12. Info. Measure of Correlation 1
H13. Info. Measure of Correlation 2


SLF1

SLF1.1 - The number of fluorescent objects in the image: Objects were identified by applying the Matlab
bwlabel function to a binarized version of the processed image. The bwlabel function defines an object as a contiguous group of non-zero pixels in an 8-connected environment (i.e., a given pixel is adjacent to each of its eight neighbors).

SLF1.2 - The Euler number of the image: The Matlab imfeature function was used to calculate the number of objects in the image minus the number of holes. A hole is defined as a contiguous group of zero-valued pixels contained entirely within an area of non-zero pixels.

SLF1.3 - The average number of above-threshold pixels per object: The mean number of non-zero pixels per object was calculated for the binarized image.

SLF1.4 - The variance of the number of above-threshold pixels per object. The variance of the number of non-zero pixels per object was also calculated.

SLF1.5 - The ratio of the size of the largest object to the smallest: This was defined as the number of pixels in the largest object divided by the number of pixels in the smallest object.

SLF1.6 - The average object distance to the cellular center of fluorescence: The center of fluorescence (COF) of the whole cell was calculated and used to determine distances to the centers of fluorescence of each object in that cell. Centers of fluorescence were calculated as:

,

where x and y are the coordinates of each pixel (in either the entire cell or a particular object), and f(x,y) is the intensity of the pixel at (x,y).

SLF1.7 - The variance of object distances from the image COF. The variance was calculated using the COF determined for SLF1.6.

SLF1.8 - The ratio of the largest to the smallest object to image COF distance: This was calculated as the distance from the image COF to the furthest object in the cell divided by the distance from the image COF to the closest object.

SLF1.9 - The fraction of the non-zero pixels in a cell that are along an edge: Edge detection was performed on each image using the Canny method () as implemented in the Matlab edge function. Canny's method calculates the gradient of the image using the derivative of a Gaussian filter. It then assigns edges to strong and weak categories. Weak edges are only included in the final output if they are connected to strong edges. This approach is less sensitive to noise in the image than other edge detection methods. The area of the binarized edge image was then divided by the area of the binarized cell image.

SLF1.10 - Measure of edge intensity homogeneity: Each image (I) was convolved separately with the kernels N and W

,

to find the intensity gradients in two orthogonal directions ( and ). The intensity of the gradient at all points in the image was calculated using

and an eight-bin histogram was calculated for the values in this edge intensity image. The final feature was calculated as the fraction of all values that fall in the first two bins of this histogram.

SLF1.11 - Measure of edge direction homogeneity 1: The overall gradient at each point in the image G was then calculated from the convolved images GN and GW calculated for SLF1.10 using

The value of each pixel in the image G is therefore the direction (from -inf; to inf;) of the intensity gradient at that point in the image, I. An eight-bin histogram was then calculated using all of the values in the gradient image G. Images with patterns containing edges oriented predominantly along a particular direction (some patterns of actin filaments, for example) result in edge gradient histograms in which a few bins will dominate. The final feature was calculated as the ratio of the largest to smallest value in the histogram.

SLF1.12 - Measure of edge direction homogeneity 2: The ratio of the largest to the next largest value in the eight-bin histogram used for SLF1.11 was calculated. This feature was included to overcome problems that may arise with values of the first measure of edge direction homogeneity becoming very large when the minimum value of the histogram is small.

SLF1.13 - Measure of edge direction difference: For the eight-bin histogram used for SLF1.11, the difference between the bins for an angle and for that angle plus p was calculated by summing bins 1 through 4 and subtracting the sum of bins 5 through 8. This difference was normalized by the sum of all eight bins.

SLF1.14 - The fraction of the convex hull area occupied by protein fluorescence: The convex hull of the protein localization image was calculated using the convhull function in Matlab and converted to a binary image. The area of the binarized protein image was then divided by the area of the convex hull image.

SLF1.15 - The roundness of the convex hull: The roundness was defined as . This value approaches 1 as the shape approaches a circle.

SLF1.16 - The eccentricity of the convex hull: The eccentricity

of the ellipse that is equivalent, based on second order moments, to the protein image convex hull was calculated using the following (from ):

where are the central moments of the protein image convex hull.


SLF2

SLF2.17 - The average object distance from the COF of the DNA image: As for SLF1.6, the distances from a reference point of objects in the protein image are calculated. However, in this case the center of fluorescence of the DNA image is used in place of the center of fluorescence of the protein image.

SLF2.18 - The variance of object distances from the DNA COF. This feature is analogous to SLF1.7 except that the DNA COF is used as the reference point.

SLF2.19 - The ratio of the largest to the smallest object to DNA COF distance. This feature is analogous to SLF1.8 except that the DNA COF is used as the reference point.

SLF2.20 - The distance between the protein COF and the DNA COF: The distance between the COF of a protein image and its corresponding DNA image is calculated.

SLF2.21 - The ratio of the area occupied by protein to that occupied by DNA: The number of pixels in the binarized protein image is divided by the number of pixels in the binarized DNA image.

SLF2.22 - The fraction of the protein fluorescence that co-localizes with DNA: The fraction of pixels in the binarized protein image that overlap with pixels in the binarized DNA image is calculated.


SLF7: SLF7.80-84 were defined based on the morphological skeleton of objects obtained by thinning using a homotopic interval.

SLF7.79 - The fraction of cellular fluorescence not included in objects: This was added to measure the amount of fluorescence that is not contained in fluorescent objects.

SLF7.80 - The average length of the morphological skeleton of objects.

SLF7.81 - The ratio of object skeleton length to the area of the convex hull of the skeleton, averaged over all objects.

SLF7.82 - The fraction of object pixels contained within the skeleton, averaged over all objects.

SLF7.83 - The fraction of object fluorescence contained within its skeleton, averaged over all objects.

SLF7.84 - The ratio of the number of branch points in skeleton to length of skeleton, averaged over all objects: A point was defined as a branch point if 3 or more of its neighbors were contained within the skeleton.


wav - Wavelet features. A set of 30 wavelet features.


SLF31: SLF31.1-18 are parameter free threshold adjacency statistics.

SLF31.1 - pftas:center_0

SLF31.2 - pftas:center_1

SLF31.3 - pftas:center_2

SLF31.4 - pftas:center_3

SLF31.5 - pftas:center_4

SLF31.6 - pftas:center_5

SLF31.7 - pftas:center_6

SLF31.8 - pftas:center_7

SLF31.9 - pftas:center_8

SLF31.10 - npftas:center_0

SLF31.11 - npftas:center_1

SLF31.12 - npftas:center_2

SLF31.13 - npftas:center_3

SLF31.14 - npftas:center_4

SLF31.15 - npftas:center_5

SLF31.16 - npftas:center_6

SLF31.17 - npftas:center_7

SLF31.18 - npftas:center_8


SLF33: SLF33.126-161 are parameter free threshold adjacency statistics calculated at mean and mean-margin instead of center.

SLF33.126 - pftas:mu_margin_0

SLF33.127 - pftas:mu_margin_1

SLF33.128 - pftas:mu_margin_2

SLF33.129 - pftas:mu_margin_3

SLF33.130 - pftas:mu_margin_4

SLF33.131 - pftas:mu_margin_5

SLF33.132 - pftas:mu_margin_6

SLF33.133 - pftas:mu_margin_7

SLF33.134 - pftas:mu_margin_8

SLF33.135 - npftas:mu_margin_0

SLF33.136 - npftas:mu_margin_1

SLF33.137 - npftas:mu_margin_2

SLF33.138 - npftas:mu_margin_3

SLF33.139 - npftas:mu_margin_4

SLF33.140 - npftas:mu_margin_5

SLF33.141 - npftas:mu_margin_6

SLF33.142 - npftas:mu_margin_7

SLF33.143 - npftas:mu_margin_8

SLF33.144 - pftas:mu_0

SLF33.145 - pftas:mu_1

SLF33.146 - pftas:mu_2

SLF33.147 - pftas:mu_3

SLF33.148 - pftas:mu_4

SLF33.149 - pftas:mu_5

SLF33.150 - pftas:mu_6

SLF33.151 - pftas:mu_7

SLF33.152 - pftas:mu_8

SLF33.153 - npftas:mu_0

SLF33.154 - npftas:mu_1

SLF33.155 - npftas:mu_2

SLF33.156 - npftas:mu_3

SLF33.157 - npftas:mu_4

SLF33.158 - npftas:mu_5

SLF33.159 - npftas:mu_6

SLF33.160 - npftas:mu_7

SLF33.161 - npftas:mu_8


SLF34: SLF34.1-10 are overlap features

SLF34.164 - overlap:prot-to-ref-overlap

SLF34.165 - overlap:fraction-above-thresh-prot-in-above-thresh-ref

SLF34.166 - overlap:fraction-of-protein-in-above-thresh-ref

SLF34.167 - overlap:fraction-of-proc-protein-in-above-thresh-ref

SLF34.168 - overlap:fraction-above-thresh-ref-in-above-thresh-prot

SLF34.169 - overlap:correlation:binprot-binref

SLF34.170 - overlap:correlation:prot-binref

SLF34.171 - overlap:correlation:prot-ref

SLF34.172 - overlap:median-prot-dist-ref

SLF34.173 - overlap:mean-prot-dist-ref


3D SLF


SLF9

SLF9.1 - The number of fluorescent objects in the image: A 3D object is defined as a group of contiguous, above-threshold voxels in a 26-connected environment.

SLF9.2 - The Euler number of the image: This is the difference between number of objects and number of holes in the image.

SLF9.3 - The average object volume: The volume of an object is defined as the number of voxels in the object.

SLF9.4 - The standard deviation of object volumes.

SLF9.5 - The ratio of the max object volume to min object volume.

SLF9.6 - The average object distance to the protein COF.

SLF9.7 - The standard deviation of object distances from the protein COF.

SLF9.8 - The ratio of the largest to the smallest object to protein COF distance.

SLF9.9 - The average object distance to the COF of the DNA image.

SLF9.10 - The standard deviation of object distances from the COF of the DNA image.

SLF9.11 - The ratio of the largest to the smallest object to DNA COF distance.

SLF9.12 - The distance between the protein COF and the DNA COF.

SLF9.13 - The ratio of the volume occupied by protein to that occupied by DNA.

SLF9.14 - The fraction of the protein fluorescence that co-localizes with DNA.

SLF9.15 - The average horizontal distance of objects to the protein COF.

SLF9.16 - The standard deviation of object horizontal distances from the protein COF.

SLF9.17 - The ratio of the largest to the smallest object to protein COF horizontal distance.

SLF9.18 - The average vertical distance of objects to the protein COF.

SLF9.19 - The standard deviation of object vertical distances from the protein COF.

SLF9.20 - The ratio of the largest to the smallest object to protein COF vertical distance.

SLF9.21 - The average object horizontal distance from the DNA COF.

SLF9.22 - The standard deviation of object horizontal distances from the DNA COF.

SLF9.23 - The ratio of the largest to the smallest object to DNA COF horizontal distance.

SLF9.24 - The average object vertical distance from the DNA COF.

SLF9.25 - The standard deviation of object vertical distances from the DNA COF.

SLF9.26 - The ratio of the largest to the smallest object to DNA COF vertical distance.

SLF9.27 - The horizontal distance between the protein COF and the DNA COF.

SLF9.28 - The signed vertical distance between the protein COF and the DNA COF: This was specifically defined to be a signed quantity in order to distinguish proteins distributed near the top of the cell from ones near the bottom.




27
SLFSLF IndexFeature NameFeature Description

SLF7

1SLF1.1object:number

SLF7

2SLF1.2object:EulerNumber

SLF7

3SLF1.3object_size:average

SLF7

4SLF1.4object_size:variance

SLF7

5SLF1.5object_size:ratio

SLF7

6SLF1.6object_distance:average

SLF7

7SLF1.7object_distance:variance

SLF7

8SLF1.8object_distance:ratio

SLF7

9SLF1.14convex_hull:fraction_of_overlap

SLF7

10SLF1.15convex_hull:shape_factor

SLF7

11SLF1.16convex_hull:eccentricity

SLF7

12SLF3.17Z_0_0

SLF7

13SLF3.18Z_1_1

SLF7

14SLF3.19Z_2_0

SLF7

15SLF3.20Z_2_2

SLF7

16SLF3.21Z_3_1

SLF7

17SLF3.22Z_3_3

SLF7

18SLF3.23Z_4_0

SLF7

19SLF3.24Z_4_2

SLF7

20SLF3.25Z_4_4

SLF7

21SLF3.26Z_5_1

SLF7

22SLF3.27Z_5_3

SLF7

23SLF3.28Z_5_5

SLF7

24SLF3.29Z_6_0

SLF7

25SLF3.30Z_6_2

SLF7

26SLF3.31Z_6_4

SLF7

27SLF3.32Z_6_6

SLF7

28SLF3.33Z_7_1

SLF7

29SLF3.34Z_7_3

SLF7

30SLF3.35Z_7_5

SLF7

31SLF3.36Z_7_7

SLF7

32SLF3.37Z_8_0

SLF7

33SLF3.38Z_8_2

SLF7

34SLF3.39Z_8_4

SLF7

35SLF3.40Z_8_6

SLF7

36SLF3.41Z_8_8

SLF7

37SLF3.42Z_9_1

SLF7

38SLF3.43Z_9_3

SLF7

39SLF3.44Z_9_5

SLF7

40SLF3.45Z_9_7

SLF7

41SLF3.46Z_9_9

SLF7

42SLF3.47Z_10_0

SLF7

43SLF3.48Z_10_2

SLF7

44SLF3.49Z_10_4

SLF7

45SLF3.50Z_10_6

SLF7

46SLF3.51Z_10_8

SLF7

47SLF3.52Z_10_10

SLF7

48SLF3.53Z_11_1

SLF7

49SLF3.54Z_11_3

SLF7

50SLF3.55Z_11_5

SLF7

51SLF3.56Z_11_7

SLF7

52SLF3.57Z_11_9

SLF7

53SLF3.58Z_11_11

SLF7

54SLF3.59Z_12_0

SLF7

55SLF3.60Z_12_2

SLF7

56SLF3.61Z_12_4

SLF7

57SLF3.62Z_12_6

SLF7

58SLF3.63Z_12_8

SLF7

59SLF3.64Z_12_10

SLF7

60SLF3.65Z_12_12

SLF7

61SLF3.66haralick:angular_second_moment

SLF7

62SLF3.67haralick:contrast

SLF7

63SLF3.68haralick:correlation

SLF7

64SLF3.69haralick:sum_of_squares

SLF7

65SLF3.70haralick:inverse_diff_moment

SLF7

66SLF3.71haralick:sum_avg

SLF7

67SLF3.72haralick:sum_var

SLF7

68SLF3.73haralick:sum_entropy

SLF7

69SLF3.74haralick:entropy

SLF7

70SLF3.75haralick:diff_var

SLF7

71SLF3.76haralick:diff_entropy

SLF7

72SLF3.77haralick:info_measure_corr_1

SLF7

73SLF3.78haralick:info_measure_corr_2

SLF7

74SLF1.9edges:area_fraction

SLF7

75SLF1.10edges:homogeneity

SLF7

76SLF1.11edges:direction_maxmin_ratio

SLF7

77SLF1.12edges:direction_maxnextmax_ratio

SLF7

78SLF1.13edges:direction_difference

SLF7

79SLF7.79fract_nonobj_fluor

SLF7

80SLF7.80obj_skel_len

SLF7

81SLF7.81obj_skel_hull_area_ratio

SLF7

82SLF7.82obj_skel_obj_area_ratio

SLF7

83SLF7.83obj_skel_obj_fluor_ratio

SLF7DNA

1SLF7.80obj_skel_len

SLF7DNA

2SLF7.81obj_skel_hull_area_ratio

SLF7DNA

3SLF7.82obj_skel_obj_area_ratio

SLF7DNA

4SLF7.83obj_skel_obj_fluor_ratio

SLF7DNA

5SLF7.84obj_skel_branch_per_len

SLF7DNA

6SLF7.79fract_nonobj_fluor

SLF7DNA

7SLF1.1object:number

SLF7DNA

8SLF1.2object:EulerNumber

SLF7DNA

9SLF1.3object_size:average

SLF7DNA

10SLF1.4object_size:variance

SLF7DNA

11SLF1.5object_size:ratio

SLF7DNA

12SLF1.6object_distance:average

SLF7DNA

13SLF1.7object_distance:variance

SLF7DNA

14SLF1.8object_distance:ratio

SLF7DNA

15SLF2.17DNA_object_distance:average

SLF7DNA

16SLF2.18DNA_object_distance:variance

SLF7DNA

17SLF2.19DNA_object_distance:ratio

SLF7DNA

18SLF2.20DNA/image:distance

SLF7DNA

19SLF2.21DNA/image:area_ratio

SLF7DNA

20SLF2.22DNA/image:overlap

SLF7DNA

21SLF1.14convex_hull:fraction_of_overlap

SLF7DNA

22SLF1.15convex_hull:shape_factor

SLF7DNA

23SLF1.16convex_hull:eccentricity

SLF7DNA

24SLF3.17Z_0_0

SLF7DNA

25SLF3.18Z_1_1

SLF7DNA

26SLF3.19Z_2_0

SLF7DNA

27SLF3.20Z_2_2

SLF7DNA

28SLF3.21Z_3_1

SLF7DNA

29SLF3.22Z_3_3

SLF7DNA

30SLF3.23Z_4_0

SLF7DNA

31SLF3.24Z_4_2

SLF7DNA

32SLF3.25Z_4_4

SLF7DNA

33SLF3.26Z_5_1

SLF7DNA

34SLF3.27Z_5_3

SLF7DNA

35SLF3.28Z_5_5

SLF7DNA

36SLF3.29Z_6_0

SLF7DNA

37SLF3.30Z_6_2

SLF7DNA

38SLF3.31Z_6_4

SLF7DNA

39SLF3.32Z_6_6

SLF7DNA

40SLF3.33Z_7_1

SLF7DNA

41SLF3.34Z_7_3

SLF7DNA

42SLF3.35Z_7_5

SLF7DNA

43SLF3.36Z_7_7

SLF7DNA

44SLF3.37Z_8_0

SLF7DNA

45SLF3.38Z_8_2

SLF7DNA

46SLF3.39Z_8_4

SLF7DNA

47SLF3.40Z_8_6

SLF7DNA

48SLF3.41Z_8_8

SLF7DNA

49SLF3.42Z_9_1

SLF7DNA

50SLF3.43Z_9_3

SLF7DNA

51SLF3.44Z_9_5

SLF7DNA

52SLF3.45Z_9_7

SLF7DNA

53SLF3.46Z_9_9

SLF7DNA

54SLF3.47Z_10_0

SLF7DNA

55SLF3.48Z_10_2

SLF7DNA

56SLF3.49Z_10_4

SLF7DNA

57SLF3.50Z_10_6

SLF7DNA

58SLF3.51Z_10_8

SLF7DNA

59SLF3.52Z_10_10

SLF7DNA

60SLF3.53Z_11_1

SLF7DNA

61SLF3.54Z_11_3

SLF7DNA

62SLF3.55Z_11_5

SLF7DNA

63SLF3.56Z_11_7

SLF7DNA

64SLF3.57Z_11_9

SLF7DNA

65SLF3.58Z_11_11

SLF7DNA

66SLF3.59Z_12_0

SLF7DNA

67SLF3.60Z_12_2

SLF7DNA

68SLF3.61Z_12_4

SLF7DNA

69SLF3.62Z_12_6

SLF7DNA

70SLF3.63Z_12_8

SLF7DNA

71SLF3.64Z_12_10

SLF7DNA

72SLF3.65Z_12_12

SLF7DNA

73SLF3.66haralick:angular_second_moment

SLF7DNA

74SLF3.67haralick:contrast

SLF7DNA

75SLF3.68haralick:correlation

SLF7DNA

76SLF3.69haralick:sum_of_squares

SLF7DNA

77SLF3.70haralick:inverse_diff_moment

SLF7DNA

78SLF3.71haralick:sum_avg

SLF7DNA

79SLF3.72haralick:sum_var

SLF7DNA

80SLF3.73haralick:sum_entropy

SLF7DNA

81SLF3.74haralick:entropy

SLF7DNA

82SLF3.75haralick:diff_var

SLF7DNA

83SLF3.76haralick:diff_entropy

SLF7DNA

84SLF3.77haralick:info_measure_corr_1

SLF7DNA

85SLF3.78haralick:info_measure_corr_2

SLF7DNA

86SLF1.9edges:area_fraction

SLF7DNA

87SLF1.10edges:homogeneity

SLF7DNA

88SLF1.11edges:direction_maxmin_ratio

SLF7DNA

89SLF1.12edges:direction_maxnextmax_ratio

SLF7DNA

90SLF1.13edges:direction_difference

SLF8

1SLF1.3object_size:average

SLF8

2SLF3.74haralick:entropy

SLF8

3SLF3.19Z_2_0

SLF8

4SLF7.79fract_nonobj_fluor

SLF8

5SLF3.71haralick:sum_avg

SLF8

6SLF3.76haralick:diff_entropy

SLF8

7SLF3.23Z_4_0

SLF8

8SLF1.9edges:area_fraction

SLF8

9SLF1.2object:EulerNumber

SLF8

10SLF1.6object_distance:average

SLF8

11SLF3.68haralick:correlation

SLF8

12SLF3.59Z_12_0

SLF8

13SLF1.8object_distance:ratio

SLF8

14SLF1.11edges:direction_maxmin_ratio

SLF8

15SLF3.47Z_10_0

SLF8

16SLF3.70haralick:inverse_diff_moment

SLF8

17SLF7.82obj_skel_obj_area_ratio

SLF8

18SLF1.1object:number

SLF8

19SLF3.24Z_4_2

SLF8

20SLF3.66haralick:angular_second_moment

SLF8

21SLF7.80obj_skel_len

SLF8

22SLF3.69haralick:sum_of_squares

SLF8

23SLF3.50Z_10_6

SLF8

24SLF1.5object_size:ratio

SLF8

25SLF7.84obj_skel_branch_per_len

SLF8

26SLF3.77haralick:info_measure_corr_1

SLF8

27SLF1.10edges:homogeneity

SLF8

28SLF3.73haralick:sum_entropy

SLF8

29SLF3.26Z_5_1

SLF8

30SLF3.78haralick:info_measure_corr_2

SLF8

31SLF3.72haralick:sum_var

SLF8

32SLF1.7object_distance:variance

SLF10

1SLF9.14The fraction of the protein fluorescence that co-localizes with DNA

SLF10

2SLF9.24The average object vertical distance from the DNA COF

SLF10

3SLF9.13The ratio of the volume occupied by protein to that occupied by DNA

SLF10

4SLF9.4The standard deviation of object volumes

SLF10

5SLF9.2The Euler number of the image

SLF10

6SLF9.1The number of fluorescent objects in the image

SLF10

7SLF9.15The average horizontal distance of objects to the protein COF

SLF10

8SLF9.21The average object horizontal distance from the DNA COF

SLF10

9SLF9.22The standard deviation of object horizontal distances from the DNA COF

SLF11

1SLF9.1The number of fluorescent objects in the image

SLF11

2SLF9.2The Euler number of the image

SLF11

3SLF9.3The average object volume

SLF11

4SLF9.4The standard deviation of object volumes

SLF11

5SLF9.5The ratio of the max object volume to min object volume

SLF11

6SLF9.6The average object distance to the protein center of fluorescenceCOF)

SLF11

7SLF9.7The standard deviation of object distances from the protein COF

SLF11

8SLF9.8The ratio of the largest to the smallest object to protein COF distance

SLF11

9SLF9.15The average horizontal distance of objects to the protein COF

SLF11

10SLF9.16The standard deviation of object horizontal distances from the protein COF

SLF11

11SLF9.17The ratio of the largest to the smallest object to protein COF horizontal distance

SLF11

12SLF9.18The average vertical distance of objects to the protein COF

SLF11

13SLF9.19The standard deviation of object vertical distances from the protein COF

SLF11

14SLF9.20The ratio of the largest to the smallest object to protein COF vertical distance

SLF11

15SLF11.15The fraction of above threshold pixels that are along an edge

SLF11

16SLF11.16The fraction of fluorescence in above threshold pixels that are along an edge

SLF11

17SLF11.17Average of angular second moment

SLF11

18SLF11.18Average of contrast

SLF11

19SLF11.19Average of correlation

SLF11

20SLF11.20Average of sum of squares of variance

SLF11

21SLF11.21Average of inverse difference moment

SLF11

22SLF11.22Average of sum average

SLF11

23SLF11.23Average of sum variance

SLF11

24SLF11.24Average of sum entropy

SLF11

25SLF11.25Average of entropy

SLF11

26SLF11.26Average of difference variance

SLF11

27SLF11.27Average of difference entropy

SLF11

28SLF11.28Average of info measure of correlation 1

SLF11

29SLF11.29Average of info measure of correlation 2

SLF11

30SLF11.30Range of angular second moment

SLF11

31SLF11.31Range of contrast

SLF11

32SLF11.32Range of correlation

SLF11

33SLF11.33Range of sum of squares of variance

SLF11

34SLF11.34Range of inverse difference moment

SLF11

35SLF11.35Range of sum average

SLF11

36SLF11.36Range of sum variance

SLF11

37SLF11.37Range of sum entropy

SLF11

38SLF11.38Range of entropy

SLF11

39SLF11.39Range of difference variance

SLF11

40SLF11.40Range of difference entropy

SLF11

41SLF11.41Range of info measure of correlation 1

SLF11

42SLF11.42Range of info measure of correlation 2

SLF12

1SLF1.3object_size:average

SLF12

2SLF3.74haralick:entropy

SLF12

3SLF3.19Z_2_0

SLF12

4SLF7.79fract_nonobj_fluor

SLF12

5SLF3.71haralick:sum_avg

SLF12

6SLF3.76haralick:diff_entropy

SLF12

7SLF3.23Z_4_0

SLF12

8SLF1.9edges:area_fraction

SLF13

1SLF1.3object_size:average

SLF13

2SLF3.74haralick:entropy

SLF13

3SLF2.22DNA/image:overlap

SLF13

4SLF3.19Z_2_0

SLF13

5SLF7.79fract_nonobj_fluor

SLF13

6SLF3.71haralick:sum_avg

SLF13

7SLF3.23Z_4_0

SLF13

8SLF3.76haralick:diff_entropy

SLF13

9SLF1.9edges:area_fraction

SLF13

10SLF3.68haralick:correlation

SLF13

11SLF1.6object_distance:average

SLF13

12SLF2.19DNA_object_distance:ratio

SLF13

13SLF1.11edges:direction_maxmin_ratio

SLF13

14SLF1.2object:EulerNumber

SLF13

15SLF3.37Z_8_0

SLF13

16SLF3.24Z_4_2

SLF13

17SLF7.82obj_skel_obj_area_ratio

SLF13

18SLF3.60Z_12_2

SLF13

19SLF2.21DNA/image:area_ratio

SLF13

20SLF3.70haralick:inverse_diff_moment

SLF13

21SLF1.1object:number

SLF13

22SLF3.50Z_10_6

SLF13

23SLF3.77haralick:info_measure_corr_1

SLF13

24SLF1.5object_size:ratio

SLF13

25SLF3.66haralick:angular_second_moment

SLF13

26SLF2.17DNA_object_distance:average

SLF13

27SLF7.84obj_skel_branch_per_len

SLF13

28SLF1.8object_distance:ratio

SLF13

29SLF1.10edges:homogeneity

SLF13

30SLF3.69haralick:sum_of_squares

SLF13

31SLF3.67haralick:contrast

SLF14

1SLF9.1The number of fluorescent objects in the image

SLF14

2SLF9.2The Euler number of the image

SLF14

3SLF9.3The average object volume

SLF14

4SLF9.4The standard deviation of object volumes

SLF14

5SLF9.5The ratio of the max object volume to min object volume

SLF14

6SLF9.6The average object distance to the protein center of fluorescenceCOF)

SLF14

7SLF9.7The standard deviation of object distances from the protein COF

SLF14

8SLF9.8The ratio of the largest to the smallest object to protein COF distance

SLF14

9SLF9.15The average horizontal distance of objects to the protein COF

SLF14

10SLF9.16The standard deviation of object horizontal distances from the protein COF

SLF14

11SLF9.17The ratio of the largest to the smallest object to protein COF horizontal distance

SLF14

12SLF9.18The average vertical distance of objects to the protein COF

SLF14

13SLF9.19The standard deviation of object vertical distances from the protein COF

SLF14

14SLF9.20The ratio of the largest to the smallest object to protein COF vertical distance

SLF21

1SLF7.80obj_skel_len

SLF21

2SLF7.81obj_skel_hull_area_ratio

SLF21

3SLF7.82obj_skel_obj_area_ratio

SLF21

4SLF7.83obj_skel_obj_fluor_ratio

SLF21

5SLF7.84obj_skel_branch_per_len

SLF21

6SLF1.3object_size:average

SLF21

7SLF1.4object_size:variance

SLF21

8SLF1.5object_size:ratio

SLF21

9SLF3.66haralick:angular_second_moment

SLF21

10SLF3.67haralick:contrast

SLF21

11SLF3.68haralick:correlation

SLF21

12SLF3.69haralick:sum_of_squares

SLF21

13SLF3.70haralick:inverse_diff_moment

SLF21

14SLF3.71haralick:sum_avg

SLF21

15SLF3.72haralick:sum_var

SLF21

16SLF3.73haralick:sum_entropy

SLF21

17SLF3.74haralick:entropy

SLF21

18SLF3.75haralick:diff_var

SLF21

19SLF3.76haralick:diff_entropy

SLF21

20SLF3.77haralick:info_measure_corr_1

SLF21

21SLF3.78haralick:info_measure_corr_2

SLF21

22SLF1.9edges:area_fraction

SLF21

23SLF1.10edges:homogeneity

SLF21

24SLF1.11edges:direction_maxmin_ratio

SLF21

25SLF1.12edges:direction_maxnextmax_ratio

SLF21

26SLF1.13edges:direction_difference

SLF27

1SLF27.1object:number

SLF27

2SLF27.2object:EulerNumber

SLF27

3SLF27.3object_size:average

SLF27

4SLF27.4object_size:variance

SLF27

5SLF27.5object_size:ratio

SLF27

6SLF27.6object_distance:average

SLF27

7SLF27.7object_distance:variance

SLF27

8SLF27.8object_distance:ratio

SLF27

9SLF27.9edges:area_fraction

SLF27

10SLF27.10edges:homogeneity

SLF27

11SLF27.11edges:direction_maxmin_ratio

SLF27

12SLF27.12edges:direction_maxnextmax_ratio

SLF27

13SLF27.13edges:direction_difference

SLF27

14SLF27.14convex_hull:fraction_of_overlap

SLF27

15SLF27.15convex_hull:shape_factor

SLF27

16SLF27.16convex_hull:eccentricity

SLF27

17SLF27.17Z_0_0

SLF27

18SLF27.18Z_1_1

SLF27

19SLF27.19Z_2_0

SLF27

20SLF27.20Z_2_2

SLF27

21SLF27.21Z_3_1

SLF27

22SLF27.22Z_3_3

SLF27

23SLF27.23Z_4_0

SLF27

24SLF27.24Z_4_2

SLF27

25SLF27.25Z_4_4

SLF27

26SLF27.26Z_5_1

SLF27

27SLF27.27Z_5_3

SLF27

28SLF27.28Z_5_5

SLF27

29SLF27.29Z_6_0

SLF27

30SLF27.30Z_6_2

SLF27

31SLF27.31Z_6_4

SLF27

32SLF27.32Z_6_6

SLF27

33SLF27.33Z_7_1

SLF27

34SLF27.34Z_7_3

SLF27

35SLF27.35Z_7_5

SLF27

36SLF27.36Z_7_7

SLF27

37SLF27.37Z_8_0

SLF27

38SLF27.38Z_8_2

SLF27

39SLF27.39Z_8_4

SLF27

40SLF27.40Z_8_6

SLF27

41SLF27.41Z_8_8

SLF27

42SLF27.42Z_9_1

SLF27

43SLF27.43Z_9_3

SLF27

44SLF27.44Z_9_5

SLF27

45SLF27.45Z_9_7

SLF27

46SLF27.46Z_9_9

SLF27

47SLF27.47Z_10_0

SLF27

48SLF27.48Z_10_2

SLF27

49SLF27.49Z_10_4

SLF27

50SLF27.50Z_10_6

SLF27

51SLF27.51Z_10_8

SLF27

52SLF27.52Z_10_10

SLF27

53SLF27.53Z_11_1

SLF27

54SLF27.54Z_11_3

SLF27

55SLF27.55Z_11_5

SLF27

56SLF27.56Z_11_7

SLF27

57SLF27.57Z_11_9

SLF27

58SLF27.58Z_11_11

SLF27

59SLF27.59Z_12_0

SLF27

60SLF27.60Z_12_2

SLF27

61SLF27.61Z_12_4

SLF27

62SLF27.62Z_12_6

SLF27

63SLF27.63Z_12_8

SLF27

64SLF27.64Z_12_10

SLF27

65SLF27.65Z_12_12

SLF27

66SLF27.66angular_second_moment

SLF27

67SLF27.67contrast

SLF27

68SLF27.68correlation

SLF27

69SLF27.69sum_of_squares

SLF27

70SLF27.70inverse_diff_moment

SLF27

71SLF27.71sum_avg

SLF27

72SLF27.72sum_var

SLF27

73SLF27.73sum_entropy

SLF27

74SLF27.74entropy

SLF27

75SLF27.75diff_var

SLF27

76SLF27.76diff_entropy

SLF27

77SLF27.77info_measure_corr_1

SLF27

78SLF27.78info_measure_corr_2

SLF27

79SLF27.79fract_nonobj_fluor

SLF27

80SLF27.80obj_skel_len

SLF27

81SLF27.81obj_skel_hull_area_ratio

SLF27

82SLF27.82obj_skel_obj_area_ratio

SLF27

83SLF27.83obj_skel_obj_fluor_ratio

SLF27

84SLF27.84obj_skel_branch_per_len

SLF30

1SLF27.1object:number

SLF30

2SLF27.2object:EulerNumber

SLF30

3SLF27.3object_size:average

SLF30

4SLF27.4object_size:variance

SLF30

5SLF27.5object_size:ratio

SLF30

6SLF27.6object_distance:average

SLF30

7SLF27.7object_distance:variance

SLF30

8SLF27.8object_distance:ratio

SLF30

9SLF27.9edges:area_fraction

SLF30

10SLF27.10edges:homogeneity

SLF30

11SLF27.11edges:direction_maxmin_ratio

SLF30

12SLF27.12edges:direction_maxnextmax_ratio

SLF30

13SLF27.13edges:direction_difference

SLF30

14SLF27.14convex_hull:fraction_of_overlap

SLF30

15SLF27.15convex_hull:shape_factor

SLF30

16SLF27.16convex_hull:eccentricity

SLF30

17SLF27.17Z_0_0

SLF30

18SLF27.18Z_1_1

SLF30

19SLF27.19Z_2_0

SLF30

20SLF27.20Z_2_2

SLF30

21SLF27.21Z_3_1

SLF30

22SLF27.22Z_3_3

SLF30

23SLF27.23Z_4_0

SLF30

24SLF27.24Z_4_2

SLF30

25SLF27.25Z_4_4

SLF30

26SLF27.26Z_5_1

SLF30

27SLF27.27Z_5_3

SLF30

28SLF27.28Z_5_5

SLF30

29SLF27.29Z_6_0

SLF30

30SLF27.30Z_6_2

SLF30

31SLF27.31Z_6_4

SLF30

32SLF27.32Z_6_6

SLF30

33SLF27.33Z_7_1

SLF30

34SLF27.34Z_7_3

SLF30

35SLF27.35Z_7_5

SLF30

36SLF27.36Z_7_7

SLF30

37SLF27.37Z_8_0

SLF30

38SLF27.38Z_8_2

SLF30

39SLF27.39Z_8_4

SLF30

40SLF27.40Z_8_6

SLF30

41SLF27.41Z_8_8

SLF30

42SLF27.42Z_9_1

SLF30

43SLF27.43Z_9_3

SLF30

44SLF27.44Z_9_5

SLF30

45SLF27.45Z_9_7

SLF30

46SLF27.46Z_9_9

SLF30

47SLF27.47Z_10_0

SLF30

48SLF27.48Z_10_2

SLF30

49SLF27.49Z_10_4

SLF30

50SLF27.50Z_10_6

SLF30

51SLF27.51Z_10_8

SLF30

52SLF27.52Z_10_10

SLF30

53SLF27.53Z_11_1

SLF30

54SLF27.54Z_11_3

SLF30

55SLF27.55Z_11_5

SLF30

56SLF27.56Z_11_7

SLF30

57SLF27.57Z_11_9

SLF30

58SLF27.58Z_11_11

SLF30

59SLF27.59Z_12_0

SLF30

60SLF27.60Z_12_2

SLF30

61SLF27.61Z_12_4

SLF30

62SLF27.62Z_12_6

SLF30

63SLF27.63Z_12_8

SLF30

64SLF27.64Z_12_10

SLF30

65SLF27.65Z_12_12

SLF30

66SLF27.66angular_second_moment

SLF30

67SLF27.67contrast

SLF30

68SLF27.68correlation

SLF30

69SLF27.69sum_of_squares

SLF30

70SLF27.70inverse_diff_moment

SLF30

71SLF27.71sum_avg

SLF30

72SLF27.72sum_var

SLF30

73SLF27.73sum_entropy

SLF30

74SLF27.74entropy

SLF30

75SLF27.75diff_var

SLF30

76SLF27.76diff_entropy

SLF30

77SLF27.77info_measure_corr_1

SLF30

78SLF27.78info_measure_corr_2

SLF30

79SLF27.79fract_nonobj_fluor

SLF30

80SLF27.80obj_skel_len

SLF30

81SLF27.81obj_skel_hull_area_ratio

SLF30

82SLF27.82obj_skel_obj_area_ratio

SLF30

83SLF27.83obj_skel_obj_fluor_ratio

SLF30

84SLF27.84obj_skel_branch_per_len

SLF30

85SLF27.85DNA_object_distance:average

SLF30

86SLF27.86DNA_object_distance:variance

SLF30

87SLF27.87DNA_object_distance:ratio

SLF30

88SLF27.88DNA/image:distance

SLF30

89SLF27.89DNA/image:area_ratio

SLF30

90SLF27.90DNA/image:overlap

SLF31

1SLF31.1pftas:center_0

SLF31

2SLF31.2pftas:center_1

SLF31

3SLF31.3pftas:center_2

SLF31

4SLF31.4pftas:center_3

SLF31

5SLF31.5pftas:center_4

SLF31

6SLF31.6pftas:center_5

SLF31

7SLF31.7pftas:center_6

SLF31

8SLF31.8pftas:center_7

SLF31

9SLF31.9pftas:center_8

SLF31

10SLF31.10npftas:center_0

SLF31

11SLF31.11npftas:center_1

SLF31

12SLF31.12npftas:center_2

SLF31

13SLF31.13npftas:center_3

SLF31

14SLF31.14npftas:center_4

SLF31

15SLF31.15npftas:center_5

SLF31

16SLF31.16npftas:center_6

SLF31

17SLF31.17npftas:center_7

SLF31

18SLF31.18npftas:center_8

SLF32

1SLF27.1object:number

SLF32

2SLF27.2object:EulerNumber

SLF32

3SLF27.3object_size:average

SLF32

4SLF27.4object_size:variance

SLF32

5SLF27.5object_size:ratio

SLF32

6SLF27.6object_distance:average

SLF32

7SLF27.7object_distance:variance

SLF32

8SLF27.8object_distance:ratio

SLF32

9SLF27.9edges:area_fraction

SLF32

10SLF27.10edges:homogeneity

SLF32

11SLF27.11edges:direction_maxmin_ratio

SLF32

12SLF27.12edges:direction_maxnextmax_ratio

SLF32

13SLF27.13edges:direction_difference

SLF32

14SLF27.14convex_hull:fraction_of_overlap

SLF32

15SLF27.15convex_hull:shape_factor

SLF32

16SLF27.16convex_hull:eccentricity

SLF32

17SLF27.17Z_0_0

SLF32

18SLF27.18Z_1_1

SLF32

19SLF27.19Z_2_0

SLF32

20SLF27.20Z_2_2

SLF32

21SLF27.21Z_3_1

SLF32

22SLF27.22Z_3_3

SLF32

23SLF27.23Z_4_0

SLF32

24SLF27.24Z_4_2

SLF32

25SLF27.25Z_4_4

SLF32

26SLF27.26Z_5_1

SLF32

27SLF27.27Z_5_3

SLF32

28SLF27.28Z_5_5

SLF32

29SLF27.29Z_6_0

SLF32

30SLF27.30Z_6_2

SLF32

31SLF27.31Z_6_4

SLF32

32SLF27.32Z_6_6

SLF32

33SLF27.33Z_7_1

SLF32

34SLF27.34Z_7_3

SLF32

35SLF27.35Z_7_5

SLF32

36SLF27.36Z_7_7

SLF32

37SLF27.37Z_8_0

SLF32

38SLF27.38Z_8_2

SLF32

39SLF27.39Z_8_4

SLF32

40SLF27.40Z_8_6

SLF32

41SLF27.41Z_8_8

SLF32

42SLF27.42Z_9_1

SLF32

43SLF27.43Z_9_3

SLF32

44SLF27.44Z_9_5

SLF32

45SLF27.45Z_9_7

SLF32

46SLF27.46Z_9_9

SLF32

47SLF27.47Z_10_0

SLF32

48SLF27.48Z_10_2

SLF32

49SLF27.49Z_10_4

SLF32

50SLF27.50Z_10_6

SLF32

51SLF27.51Z_10_8

SLF32

52SLF27.52Z_10_10

SLF32

53SLF27.53Z_11_1

SLF32

54SLF27.54Z_11_3

SLF32

55SLF27.55Z_11_5

SLF32

56SLF27.56Z_11_7

SLF32

57SLF27.57Z_11_9

SLF32

58SLF27.58Z_11_11

SLF32

59SLF27.59Z_12_0

SLF32

60SLF27.60Z_12_2

SLF32

61SLF27.61Z_12_4

SLF32

62SLF27.62Z_12_6

SLF32

63SLF27.63Z_12_8

SLF32

64SLF27.64Z_12_10

SLF32

65SLF27.65Z_12_12

SLF32

66SLF27.66angular_second_moment

SLF32

67SLF27.67contrast

SLF32

68SLF27.68correlation

SLF32

69SLF27.69sum_of_squares

SLF32

70SLF27.70inverse_diff_moment

SLF32

71SLF27.71sum_avg

SLF32

72SLF27.72sum_var

SLF32

73SLF27.73sum_entropy

SLF32

74SLF27.74entropy

SLF32

75SLF27.75diff_var

SLF32

76SLF27.76diff_entropy

SLF32

77SLF27.77info_measure_corr_1

SLF32

78SLF27.78info_measure_corr_2

SLF32

79SLF27.79fract_nonobj_fluor

SLF32

80SLF27.80obj_skel_len

SLF32

81SLF27.81obj_skel_hull_area_ratio

SLF32

82SLF27.82obj_skel_obj_area_ratio

SLF32

83SLF27.83obj_skel_obj_fluor_ratio

SLF32

84SLF27.84obj_skel_branch_per_len

SLF32

85SLF27.85DNA_object_distance:average

SLF32

86SLF27.86DNA_object_distance:variance

SLF32

87SLF27.87DNA_object_distance:ratio

SLF32

88SLF27.88DNA/image:distance

SLF32

89SLF27.89DNA/image:area_ratio

SLF32

90SLF27.90DNA/image:overlap

SLF32

91SLF31.1pftas:center_0

SLF32

92SLF31.2pftas:center_1

SLF32

93SLF31.3pftas:center_2

SLF32

94SLF31.4pftas:center_3

SLF32

95SLF31.5pftas:center_4

SLF32

96SLF31.6pftas:center_5

SLF32

97SLF31.7pftas:center_6

SLF32

98SLF31.8pftas:center_7

SLF32

99SLF31.9pftas:center_8

SLF32

100SLF31.10npftas:center_0

SLF32

101SLF31.11npftas:center_1

SLF32

102SLF31.12npftas:center_2

SLF32

103SLF31.13npftas:center_3

SLF32

104SLF31.14npftas:center_4

SLF32

105SLF31.15npftas:center_5

SLF32

106SLF31.16npftas:center_6

SLF32

107SLF31.17npftas:center_7

SLF32

108SLF31.18npftas:center_8

SLF33

1SLF27.66haralick:angular_second_moment

SLF33

2SLF27.67haralick:contrast

SLF33

3SLF27.68haralick:correlation

SLF33

4SLF27.69haralick:sum_of_squares

SLF33

5SLF27.70haralick:inverse_diff_moment

SLF33

6SLF27.71haralick:sum_avg

SLF33

7SLF27.72haralick:sum_var

SLF33

8SLF27.73haralick:sum_entropy

SLF33

9SLF27.74haralick:entropy

SLF33

10SLF27.75haralick:diff_var

SLF33

11SLF27.76haralick:diff_entropy

SLF33

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SLF34

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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SLF35

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43SLF33.159npftas:mu_6

SLF35

44SLF34.164overlap:prot-to-ref-overlap

SLF35

45SLF34.165overlap:fraction-above-thresh-prot-in-above-thresh-ref

SLF35

46SLF34.169overlap:correlation:binprot-binref

SLF35

47SLF34.170overlap:correlation:prot-binref

SLF35

48SLF34.171overlap:correlation:prot-ref

SLF35

49SLF34.173overlap:mean-prot-dist-ref

SLF36

1SLF3.67haralick:contrast

SLF36

2SLF3.75haralick:diff_var

SLF36

3SLF33.38contrast-after-dsample-1

SLF36

4SLF33.39correlation-after-dsample-1

SLF36

5SLF33.75info_measure_corr_2-after-dsample-3

SLF36

6SLF33.79sum_of_squares-after-dsample-4

SLF36

7SLF33.80inverse_diff_moment-after-dsample-4

SLF36

8SLF33.82sum_var-after-dsample-4

SLF36

9SLF33.84entropy-after-dsample-4

SLF36

10SLF33.90contrast-after-dsample-5

SLF36

11SLF33.91correlation-after-dsample-5

SLF36

12SLF33.95sum_var-after-dsample-5

SLF36

13SLF33.99diff_entropy-after-dsample-5

SLF36

14SLF33.100info_measure_corr_1-after-dsample-5

SLF36

15SLF33.108sum_var-after-dsample-6

SLF36

16SLF27.1object:number

SLF36

17SLF27.3object_size:average

SLF36

18SLF27.5object_size:ratio

SLF36

19SLF27.9edges:area_fraction

SLF36

20SLF27.10edges:homogeneity

SLF36

21SLF27.11edges:direction_maxmin_ratio

SLF36

22SLF27.80obj_skel_len

SLF36

23SLF27.81obj_skel_hull_area_ratio

SLF36

24SLF27.82obj_skel_obj_area_ratio

SLF36

25SLF27.83obj_skel_obj_fluor_ratio

SLF36

26SLF27.84obj_skel_branch_per_len

SLF36

27SLF31.2pftas:center_1

SLF36

28SLF31.5pftas:center_4

SLF36

29SLF31.8pftas:center_7

SLF36

30SLF31.9pftas:center_8

SLF36

31SLF31.11npftas:center_1

SLF36

32SLF31.13npftas:center_3

SLF36

33SLF31.18npftas:center_8

SLF36

34SLF33.144pftas:mu_0

SLF36

35SLF33.148pftas:mu_4

SLF36

36SLF33.150pftas:mu_6

SLF36

37SLF33.154npftas:mu_1

SLF36

38SLF33.156npftas:mu_3

SLF36

39SLF33.157npftas:mu_4

SLF36

40SLF33.158npftas:mu_5

SLF36

41SLF33.161npftas:mu_8

SLF36

42SLF34.167overlap:fraction-of-proc-protein-in-above-thresh-ref

SLF36

43SLF34.170overlap:correlation:prot-binref

SLF36

44SLF34.171overlap:correlation:prot-ref

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Last Updated: 16 Jan 2012




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